Skip to main content

Full text of "Zoological names. A list of phyla, classes, and orders, prepared for section F, American Association for the Advancement of Science"

See other formats


ZOOLOGICAL  NAMES 

A  List  of 
Phyla,  Classes,  and  Orders 


Prepared  for 
Section  F 

American  Association  for  the  Advancement 
of  Science 


353 

•  P42 
1949 


Durham,  North  Carolina 
1949 


ZOOLOGICAL  NAMES 


A  List  of 
Phyla,  Classes,  and  Orders 


^ 


Prepared  for 

Section  F 

American  Association  for  the  Advancement 
of  Science 


Durham,  North  Carolina 
1949 


PREFACE 

American  zoological  writers  who  have  published  books  or  scientific  papers  during  the 
past  decade  show  little  uniformity  in  their  reference  to  larger  groups  of  animals.  Of 
course  no  zoologist  wishes  to  be  limited  in  the  choice  of  names  which  seem  to  be  most 
appropriate;  no  one  wishes  to  have  imposed  on  him  terms  which  he  is  ordered  to  use;  and 
none  desires  to  have  nomeclature  become  fixed  and  inflexible.  Of  all  the  rewards  that 
Science  gives  to  her  retainers  those  most  prized  and  enjoyed  are  independence  and  the 
right  to  change  old  dogmas  as  soon  as  the  discovery  of  new  truths  makes  them  illogical. 
Yet  because  the  species  of  animals  now  number  several  hundreds  of  thousands,  the 
sorting  of  species  into  groups  must  necessarily  be  continually  more  accurate  and  critical. 
Furthermore,  there  are  men  who  through  ignorance,  inertia,  or  bigotry  take  little  or  no 
cognizance  of  the  progress  of  current  scientific  evidence  and  opinion.  For  example,  cer- 
tain writers  of  recent  textbooks  use  such  old  terms  as  Annulata  and  Anura  when  the 
most  thoughtful  and  competent  students  of  annelids  and  amphibians  agree  that  Annelida 
and  Salientia  are  to  be  preferred.  Dr.  R.  E.  Snodgrass,  who  has  recently  studied  the 
anatomy  of  onychophorans  in  detail,  is  convinced  that  these  animals  cannot  be  called 
arthropods  because  their  muscular  systems  have  quite  different  relations  to  nervous 
structures  from  those  which  obtain  in  arthropods.  Dr.  J.  P.  Moore  does  not  believe  that 
onychophorans  may  be  included  among  amielids.  There  seems  to  be  no  alternative  but 
to  set  off  Onychophora  as  a  group  by  itself. 

Those  who  know  animals  and  their  classifications  in  detail  perhaps  may  enjoy  using 
various  names  for  the  same  group  and  delight  in  thus  displaying  their  technical  knowledge, 
but  a  novice  who  refers  to  two  or  three  textbooks  and  finds  the  same  group  called  by 
two  or  three  names  is  greatly  confused.  In  an  attempt  to  bring  about  greater  uniformity 
and  more  careful  selection  in  the  use  of  zoological  group  names  the  writer  was  authorized 
to  form  an  intormal  committee  and  to  prepare  a  list  of  phyla,  classes,  and  orders  of 
animals  which  would  represent  current  good  usage.  This  action  was  taken  by  Section  F 
of  the  American  Association  for  the  Advancement  of  Science  at  the  Boston  Meeting ; 
December,  1933. 

Since  then  many  persons  who  appeared  to  be  interested  have  been  consulted.  The 
following  gave  advice  in  connection  with  the  groups  mentioned  : 

Invertebrata :  A.  H.  Clark,  B.  G.  Chitwood,  Roy  Miner,  L.  E.  Noland,  H.  S.  Pratt, 
Waldo  Schmitt,  H.  J.  Van  Cleave. 

Protozoa :  L.  E.  Noland,  and  forty-odd  protozoologists  to  whom  he  sent  a  question- 
naire. 

Porifera :  //.  V.  Wilson, 

Coelenterata  :  Elizabeth  Dekhmann  (except  Zoantharia). 

Aschelminthes  :  B.  G.  Chitwood,  M.  C.  Hall. 

Platyhehranthes  :  E.  W.  Price;  Cestoda  :  G.  R.  La  Rue;  Nemertea  :  W.  R.  Coe. 

Nemathelminthes :  B.  G.  Chitwood,  G.  Steiner,  Acanthoccphala :  //.  /.   Van   Cleave. 

Bryozoa :  R.  S.  Sassier,  R.  C.  Osburn. 

Echinodermata :  A.  H.  Clark,  H.  L.  Clark,  E.  Kirk. 

Brachiopoda  :  G.  A.  Cooper. 

Enteropneusta :  W.  E.  Ritter. 

Annelida :  J.  P.  Moore,  A.  L.  Tread  well. 

Arthropoda :  H.  E.  Ewing,  R.  E.  Snodgrass. 

Crustacea:  E.  A.  Birge,  Norma  C.  Furtos,  H.  A.  Pilsbry,  Mary  J.  Rathbun.  Waldo 
Schmitt,  J.  P.  Visscher,  C.  B.  Wilson. 

[2] 


Pycnogonidia  :  L.  J.  Cole;  Arachiiida  :  //.  £.  Eiving. 

Insecta :  C.  L.  Kennedy,  Z.  P.  Mctcalj.  J.  G.  Needham,  H.  G.  Barber,  H.  E.  Ewing. 

Mollusca :  P.  Bartsch,  H.  Rehder. 

Chordata  :  C.  W.  Gilmorc,  \V.  K.  Gregory. 

Tunicata :  IV.  E.  Ritter,  W.  G.  Van  Name. 

Pisces  :  W.  K.  Gregory,  C.  L.  Hubbs,  G.  S.  Myers. 

Amphibia  :  Helen  T.  Gaige,  G.  K.  Noble. 

Reptilia :  Helen  T.  Gaige,  G.  K.  Noble. 

Aves :  A.  Wetmore. 

Mammalia :  L.  R.  Dice,  C.  W.  Gilmore,  Remington  Kellogg. 

Those  whose  names  appear  in  italics  furnished  essentially  the  terms  that  appear  in  the 
following  pages.  While  the  writer  gladly  acknowledges  the  obligation  to  those  persons 
and  appreciates  greatly  the  work  they  did,  they  are  not  to  be  held  responsible  for  the 
published  list.  The  differences  of  opinion  among  zoologists  in  the  United  States  were 
found  to  be  so  great  that  there  was  little  hope  of  agreement  and  the  editor  finally  did 
what  seemed  to  him  best. 

Perhaps  forty  per  cent  of  those  consulted  strongly  urged  the  use  of  uniform  endings 
for  the  larger  groups  of  animals.  The  difficulty  encountered  in  this  connection  was  that 
protozoologists,  helminthologists.  conchologists,  ornitholgists,  etc.  had  already  moved 
more  or  less  in  the  direction  of  uniformity  and  each  group  of  specialists  had  a  tendency 
to  adopt  a  different  system.  After  long  consideration  the  editor  has  used  the  system  in 
vogue  among  protozoologists  and  students  of  some  other  groups,  as  it  seemed  to  be  at 
present  more  generally  in  use  than  any  other.  The  endings  are  as  follows :  Phylum  and 
Subphylum,  — a;  Class,  — ea;  Subclass,  — ia  ;  Order,  — ida ;  Suborder,  — ina.  In  presenting 
this  report  to  Section  F  it  is  not  supposed  that  uniform  endings  shall  be  adopted  as  official, 
or  even  recommended.  The  zoological  world  had  perhaps  best  be  left  to  make  its  own 
choice.  In  the  list  the  names  in  the  first  column  represent  what  the  editor  after  con- 
sultation believes  to  be  best  current  usage;  those  in  the  second  column  are  what  a  few 
zoologists  would  like  to  see  used,  but  perhaps  in  presenting  these  the  editor  is  "a  hundred 
years  ahead  of  his  time."  In  any  attempt  to  attain  uniformity  difficulties  arise.  For 
example,  Dr.  Steiner  points  out  that  two  names  suggested  as  suborders  under  Nematoda 
are  objectionable  because  identical  names  already  have  good  standing  as  genera  in  the 
same  group  (Rhabditina  Cobb  1922,  Chromadorina  Filipjev  1926). 

Another  matter  about  which  there  is  wide  disagreement  is  whether  there  should  be 
few  or  many  groups — the  old  controversy  between  "lumpers"  and  "splitters."  Shall  there 
be  ten  or  thirty  phyla,  or  orders  of  insects  ?  The  position  has  been  taken  that  it  is  more 
truthful,  and  therefore  more  scientific,  to  present  many  groups ;  otherwise  it  is  necessary 
to  compromise  to  the  extent  of  grouping  things  together  which  have  remote  or  insig- 
nificant relationships.  To  the  objection  that  students  cannot  be  expected  to  master  such 
a  long  list  of  technical  jargon  there  is  only  one  answer.  It  is  better  to  tell  students  part 
of  the  truth  frankly  than  to  falsely  affirm  that  they  are  learning  a  complete  and  simple 
story.  Biology  is  not  always  simple  and  clear.  It  therefore  seems  proper  to  say  to  a 
student  in  an  elementary  course,  "of  the  thirty-odd  phyla  of  animals,  or  orders  of  insects, 
only  ten  or  a  dozen  which  are  of  particular  biological  significance  to  you  will  be  studied," 
rather  than,  "for  convenience  and  simplicity  all  animals,  or  all  insects,  are  presented 
under  ten  phyla,  or  orders,  which  may  be  easily  remembered."  Science  is  truth. 
Classification  must  be  expected  to  grow  more  complex  as  knowledge  increases  and 
becomes  more  critical. 

In  most  groups  the  authors'  names  and  the  dates  following  correctly  indicate  when  a 
particular  scientific  name  was  given  to  a  group,  but  these  should  be  used  with  caution 
for  in  cases  where  no  specialist  could  be  found  to  furnish  accurate  information  (birds, 
fishes,  etc.),  the  editor  browsed  through  systematic  works  and  found  such  information, 

[3] 


some  of  which  later  proved  to  be  incorrect.     For  example,  some  references  credited  to 

Lamarck  by  various  writers  are  for  vernacular  names  which  should  have  no  scientific 

standing  as  scientific  names.     The  following  groups  have  been  carefully   corrected   by 

specialists  and  are  believed  to  be  reliable  for  authors  and  dates :    Protozoa,   Porifera, 

Coelenterata  (except  Zoantharia),  Platyhelminthes,  Nemertea,  Acanthocephala,  Nematoda, 

Bryozoa,  Brachiopoda,   Echinodermata,  Annelida,  Arachnida,   Crustacea,   Insecta,   Tuni- 

cata,  Amphibia,  Reptilia,  and  Mammalia ;  other  groups  may  not  be. 

A.  S.  Pearse,  Editor. 
Duke  University,  June  1,  1936. 

SECOND  EDITION 

As  there  is  continual  demand  for  this  list,  which  has  been  out  of  print  for  several 
years,  it  seems  desirable  to  publish  a  second  edition.  A  few  changes  have  been  made 
and  the  following  persons  made  suggestions  for  the  revision.  A.  H.  Clark  suggested 
the  term  Hemizoa  and  other  general  changes.  Grateful  acknowledgment  is  made  to 
G.  W.  Wharton  who  corrected  the  proofs  for  the  list.  An  excellent  discussion  of  various 
modern  systems  of  classification  will  be  found  in  Libbie  H.  Hyman's  "The  Invertebrates : 
Protozoa  through  Ctenophora,"  1940:22-43. 

Invertebrata :  A.  H.  Clark,  C.  G.  Bookhout;  Protozoa:  R.  R.  Kudo;  Porifera:  M.  W. 

de    Laubenfels ;    Nemertea :    W.    R.    Coe ;    Acanthocephala :    H.    J.    Van    Cleave ; 

Bryozoa:  R.  C.  Osburn;  Annelida:  Grace  E.  Pickford;  Arthropoda :  R.  E.  Snod- 

grass ;    Pycnogonida :    Joel    W.    Hedgpeth ;    Arachnida:    Elizabeth    B.    Bryant; 

Acarina:  G.  W.  Wharton;  Myriapoda:   Nelle  B.  Causey;  Crustacea:   Fenner  A. 

Chace;  Insecta:  C.  F.  W.  Muesebeck  and  colleagues;  Tunicata:  W.  G.  Van  Name; 

Vertebrata :  E.  C.  Horn,  I.  E.  Gray. 

T^.TT.        .       r-  ,       .,^.^  A.  S.  Pearse,  Editor. 

Duke  University,  September  1,  1946. 

THIRD  EDITION 

The  second  edition  of  this  list  was  so  full  of  errors  that  it  was  never  sent  to  the 
general  biological  public.  In  the  revision  for  this  third  edition  the  list  of  groups  that 
has  "uniform"  endings  has  been  omitted.  The  following  persons  have  furnished  informa- 
tion for  the  groups  indicated  :  Invertebrata,  Libbie  H.  Hyman  and  M.  W.  de  Laubenfels ; 
Protozoa,  R.  R.  Kudo;  Porifera,  M.  W.  de  Laubenfels;  Platyhelminthes,  Libbie  H. 
Hyman  and  H.  W.  Manter ;  Cestoda,  Robt.  P.  Dollfus;  Nemertea,  Libbie  H.  Hyman 
and  Arthur  G.  Humes ;  Brachiopoda,  Preston  E.  Cloud,  Jr. ;  Annelida,  Olga  Hartman  and 
Grace  E.  Pickford ;  Crustacea,  Fenner  A.  Chace,  Jr. ;  Arachnida,  Alexander  Petrun- 
kevitch ;  Acari,  G.  W.  Wharton ;  Pycnogonida,  Joel  W.  Hedgpeth ;  Myriapoda,  Nelle  B. 
Causey;  Insecta,  F.  M.  Carpenter  and  Charles  L.  Remington;  Mollusca,  H.  A.  Rehder; 
Tunicata,  W.  G.  Van  Name  and  A.  G.  Huntsman;  Fishes,  Carl  H.  Hubbs;  Amphibia 
and  Reptilia,  Grace  L.  Orton;  Aves,  Ernst  Mayr  and  Alexander  Wetmore.  For  trema- 
todes  Dawes  (1946)  "the  Trematoda"  has  been  followed,  for  fishes,  amphibians,  and  rep- 
tiles the  classification  in  Romer's  (1945)  "Vertebrate  Paleontology"  has  been  used,  and 
for  mammals  that  in  Simpson's  (1945)  "The  Principles  of  Classification  and  a  Classifica- 
tions of  Mammals"  has  been  followed. 

ASP 
July  1,  1948.  ■     ■     ' 

FOURTH   EDITION 

In  preparation  of  this  edition  helpful  suggestions  have  been  received  from  Austin  H. 

Clark,  Herbert  Friedmann,  J.  B.  S.  Haldane,   Harald  A.   Rehder,   Curtis  W.   Sabrosky, 

and  Alexander  Wetmore. 

ASP 
October  30,  1949. 

[4] 


A  List  of  Phyla,  Classes,  and  Orders 

Extinct  groups  are  indicated  by  an  asterisk  (*) 

Kingdom  Animalia  Linnaeus  1758 
Subkingdom  Protozoa  Goldfus  1817 
Phylum  Protozoa  Goldfus  1817 
Subphylum  Plasmodroma  Doflein  1901 
Class  Mastigophora  Diesing  1865 
Subclass  Phytomastigina  Doflein  1916 
Order  Chysomonadina  Stein  1878 

Suborder  Euchrysomonadina  Pascher  1913 
Suborder  Rhizochrysidina  Pascher  1913 
Suborder  Chrysocapsina  Pascher  1913 
Order  Cryptomonadina  Stein  1878 

Suborder  Eucryptomonadina  Pascher  1913 
Suborder  Phoecapsina  Pascher  1913 
Order  Phytomonadina  Blochmann  1895 
Order  Euglenoidina  Butschli  1884 
Order  Chloromonadina  Klebs  1892 
Order  Dinoflagellata  Butschli  1885 
Suborder  Porocentrinea  Poche  1913 
Suborder  Peridiniinea  Poche  1913 
Suborder  Cystoflagellata  Haeckel  1873 
Subclass  Zoomastigina  Doflein  1916 
Order  Rhizomastigina  Biitschli  1883 
Order  Protomonadina  Blochmann  1895 
Order  Polymastigina  Blochmann  1895 
Suborder  Monomonadina  Kudo  1939 
Suborder  Diplomonadina  Kudo  1939 
Suborder  Polymonadina  Kudo  1939 
Order  Hypermastigina  Grassi  1911 
Class  Sarcodina  Hertwig  &  Lesser  1874 
Subclass  Rhizopoda  von  Siebold  1845 
Order  Proteomyxa  Lankester  1885 
Order  Mycetozoa  de  Bary  1859 
Suborder  Eumycetozoa  Zopf  1884 
Suborder  Sorophora  Zopf  1887 
Order  Amoebina  Ehrenberg  1830 
Order  Testacea  Schultze  1854 
Order  Foraminifera  D'Orbigny  1826 
Subclass  Actinopoda  Calkins  1909 
Order  Heliozoa  Haeckel  1866 
Order  Radiolaria  J.  Miiller  1858 
Class  Sporozoa  Leuckart  1879 
Subclass  Telosporidia  Schaudinn  1900 
Order  Gregarinida  Lankester  1866 
Suborder  Eugregarinaria  Doflein  1901 
Suborder  Schizogregarinaria  Leger  1900 
Order  Coccidia  Leuckart  1879 
Suborder  Eimeridea  Leger  1911 

[S] 


Suborder  Adeleidea  Leger  1911 
Order  Haemosporidia  Danilewsky  1886 
Subclass  Acnidosporidia  Cepede  1911 
Order  Sarcosporidia  Balbiani  1882 
Order  Haplosporidia  Caullery  &  Mesnil  1899 
Subclass  Cnidosporidia  Doflein  1901 
Order  Myxosporidia  Biitschli  1881 
Suborder  Eurysporea  Kudo  1920 
Suborder  Sphaerosporea  Kudo  1920 
Suborder  Platysporea  Kudo  1920 
Order  Actinoniyxidia  Stolg  1899 
Order  Microsporidia  Balbiani  1883 

Suborder  Monocnidea  Leger  &  Hesse  1922 
Suborder  Dicnidea  Leger  &  Hesse  1922 
Order  Helicosporidia  Kudo  1931 
Subphylum  Ciliophora  Doflein  1901 
Class  Ciliata  Perty  1852 
Subclass  Protociliata  Metcalf  1918 
Subclass  Euciliata  Metcalf  1918 
Order  Holotricha  Stein  1859 

Suborder  Astomata  Schwiakoff  1896 
Suborder  Gymnostomata  Biitschli  1889 
Suborder  Trichostoniata  Biitschli  1889 
Suborder  Hymenostomata  Hickson  1903 
Suborder  Thigmotricha  Chatton  &  Lwoff  1926 
Suborder  Apostomea  Qiatton  &  Lwoff  1935 
Order  Spirotricha  Biitschli  1889 
Suborder  Heterotricha  Stein  1859 
Suborder  Oligotricha  Biitschli  1887 
Suborder  Ctenostomata  Kahl  1932 
Suborder  Hypotricha  Stein  1859 
Order  Chonotricha  Wallengren  1895 
Order  Peritricha  Stein  1859 
Suborder  Sessilia  Kahl  1935 
Suborder  Mobilia  Kahl  1935 
Class  Suctoria  Claparede  &  Lachmann  1858 
Subkingdom  Mesozoa  van  Beneden  1877 
Subkingdom  Parazoa  Sollas  1884 
Phylum  Porifera  Grant  1835 
Class  Calcispongiae  Schmidt  1862 
Order  Asconosa  de  Laubenfels  1936 
Order  Syconosa  de  Laubenfels  1936 
Class  Hyalospongiae  Vosmaer  1885 
Order  Hexasterophora  Schuize  1899 
Order  Amphidiscophora  Schuize  1899 
Class  Desmospongiae  Sollas  1875 
Order  Carnosa  Carter  1875 
Order  Choristida  Sollas  1888 
Order  Epipolasida  Sollas  1888 
Order  Hadromerina  Topsent  1898 
Order  Poecilosclerina  Topsent  1898 
Order  Haplosclerina  Topsent  1898 

[6] 


Order  Keratosa  Grant  1861 
Phylum  *Archaeocyatha  Borneniann  1884 
Subkingdom  Metazoa  Haeckel  1877 
Branch  Enterozoa  Lankcster  1878 
Superseries  Radiata  Schimkewitsch  1891 
Phylum  Coelenterata  Leuckart  1848 
Class  Hydrozoa  Huxley  1857 
Order  Hydroida  Huxley  1857 
Order  Trachylina  Haeckel  1877 
Order  Milleporina  Hickson  1899 
Order  Stylasterina  Hickson  1899 
Order  Siphonophora  Esclischoltz  1829 
Suborder  Calycophorae  Leuckart  1854 
Suborder  Rhizophysaliae  Chun  1882 
Suborder  Chondrophorae  Chamisso  &  Eysenhard  1821 
Order  *Graptolitoidea  Lapworth  1873 

Suborder  *Monoprionidea  Hopkinson  1875 
Suborder  *Diprionidea  Hopkinson  1875 
Suborder  *Retiolithidea  Lapworth  1873 
Class  Scyphozoa  Grobben  1910 

Order  Lucernariidea  Johnston  1847 
Order  Charybdeidea  Gegenbaur  1856 
Order  Corona  Vanhoffen  1892 
Order  Semaeostonieae  L.  Agassiz  1862 
Order  Rhizostomeae  Cuvier  1799 
Class  *Stromatoporoidea  Nicholson  &  Murie  1878 
Class  Anthozoa  Ehrenberg  1834 
Subclass  Alcyonaria  Mihie-Edwards  &  Haime  1857 
Order  Stolonifera  Hickson  1883 
Order  Telestacea  Hickson  1930 
Order  Alcyonacea  Verrill  1866 
Order  Gorgonacea  Verrill  1866 

Suborder  Scleraxonia  Wright  &  Struder  1889 
Suborder  Holaxonia  Wright  &  Struder  1889 
Order  Coenothecalia  Bourne  1900 
Order  Pennatulacea  Verrill  1866 
Subclass  Zoantharia  Blainville  1830 
Order  Cerianthidea  M.-Edw.  &  Haime  1857 
Order  Antipitharia  Dana  1846 
Order  Zoanthidea  Dana  1846 
Order  Edwardsiidea  R.  Hertwig  1888 
Order  Proactiniae  McMurrich  1892 
Order  Actinaria  McMurrich  1829 

Suborder  Ptychodaetiaria  Stephenson  1921 
Suborder  Coralliomorpharia  Carlgren  1936 
Suborder  Actiniaria  Carlgren  1925 
Order  Madreporaria  Linnaeus  1748 

Suborder  Aporosa  M.-Edw.  &  Haime  1857 
Suborder  Fungacea  M.-Edw.  &  Haime  1857 
Suborder  Perforata  M.-Edw.  &  Haime  1857 
Phylum  Ctenophorae  Eschscholtz  1829 
Qass  Tentaculata  Wagner  1834 

[7] 


Order  Cydippidea  Lesson  1843 

Order  Lobata  Eschscholtz  1829 

Order  Cestoidea  Lesson  1843 

Order  Platyctenea  Bourne  1900 
Class  Nuda  Bourne  190O 

Order  Beroidea  Eschscholtz  1829 
Subbranch  Triploblastica  Lankester  1900 
Superseries  Bilateria  Schimkewitsch  1891 
Series  Enterocoela  Lankester  1876 
Subseries  Acoelomata  Schimkewitsch  1891 
Phylum  Platyhelminthes  Gegenbaur  1859 
Class  Turbellaria  Ehrenberg  1831 

Order  Acoela  Uljanin  1870 

Order  Rhabdocoela  Ehrenberg  1831 

Suborder  Temnocephalida  Haswell  1888 

Order  Alloiocoela  Graff  1882  (Alloeocoela  Spengel  1884) 

Order  Tricladida  (Lang  1881) 

Order  Polycladida  (Lang  1881)  Graff  1885 
Class  Trematoda  Rudolphi  1808 
Subclass  Monogenea  Carus  1863 

Order  Monopisthocotylea  Odhner  1912 

Order  Polyopisthocotylea  Odhner  1912 
Subclass  Aspidogastrea  (Burmeister  1856)  Monticelli  1892 
Subclass  Digenea  Carus  1863 

Order  Gasterostoma  Odhner  1905 

Order  Prosostomata  Odhner  1905 
Class  Cestoidea  Rudolphi  1808 
Subclass  Cestodaria  Monticelli  1892 

Order  Amphilinidea  Poche  1922 

Order  Gyrocotylidea  Benham  1901 
Subclass  Cestoda  van  Beneden  1848 

Order  Pseudophyllidea  Carus  1863 

Order  Cyclophyllidea  Braun  1900 

Order  Tetraphyllidea  Braun  1900 

Order  Trypanorhyncha  Diesing  1863 

Order  Heterophyllidea  Southwell  1925 
Phylum  Nemertea  Delle  Chiaje  1841 
Class  Anopla  Schultze  1852 

Order  Paleonemertea  Hubrecht  1879 

Order  Heteronemertea  Burger  1892 
Qass  Enopla  Schultze  1852 

Order  Hoplonemertea  Hubrecht  1879 
Suborder  Monostilifera  Brinkmann  1917 
Suborder  Polystilifera  Brinkmann  1917 

Order  Bellonemertea  Coe  1905 
Phylum  Acanthocephala  Rudolphi  1808 
Class  Eoacanthocephala  Van  Cleave  1936 

Order  Gyracanthocephala  Van  Cleave  1936 

Order  Neoacanthocephala  Van  Cleave  1936 
Class  Metacanthocephala  Van  Cleave  1948 

Order  Palaeacanthocephala  Meyer  1931 

Order  Archiacanthocephala  Meyer  1931 

[8] 


Subseries  Pseudocoelomata  Schimkewtsch  1891 
Superphylum  Aschelminthes  Grobben  1909 
Phylum  Rotatoria  Ehrenberg  1838 
Class  Seisonidea  Wesenberg-Lund  1899 
Class  Bdelloidea  Hudson  &  Gosse  1889 
Class  Monogononta  Ehrenberg  1832 
Order  Ploima  Hudson  &  Gosse  1889 
Order  Flosculariacea  Harring  1913 
Order  Collothecacea  Harring  1913 
Phylum  Gastrotricha  Metchnikof  1864 
Order  Macrodasyoidea  Remane  1926 
Order  Chaetonotoidea  Remane  1926 
Phylum  Echinodera  Zehnka  1928  (Kinorhyncha  Reinhard  1887) 
Phylum  Nematomorpha  Vejovsky  1866 
Class  Gordididea  Orthepp  1924 
Class  Nectonematoidea  Rauther  1930 
Phylum  Nematoidea  Rudolphi  1808 

Class  Phasmidia  Qiitwood  &  Chitwood  1933 
Order  Rhabditida  Chitwood  1933 
Suborder  Rhabditata  Chitwood  1933 
Suborder  Ascaridata  Skrjabin  1915 
Suborder  Strongylata  Railliet  &  Henry  1913 
Order  Spirurida  Chitwood  1933 

Suborder  Spirurata  Railhet  &  Henry  1913 
Suborder  Camallanta  Chitwood  1936 
Class  Aphasmidia  Chitwood  &  Chitwood  1933 
Order  Enoplata  Chitwood  1933 
Suborder  Enoplata  Filipjev  1929 
Suborder  Dorylaimata  Chitwood  1933 
Suborder  Dioctophymata  Skrjabin  1923 
Order  Chromadorida  Chitwood  1933 
Suborder  Monohysterata  FiHpjev  1929 
Suborder  Desmoscolecata  Filipjev  1929 
Suborder  Chromadorata  Filipjev  1929 
Phylum  Tardigrada  Spallanzani  1726 
Class  Heterotardigrada  Marcus  1927 
Order  Arthrotardigrada  Marcus  1927 
Order  Echiniscoidea  Richters  1926 
Class  Eutardigrada  Richters  1926 
Subseries  Eucoelomata  Schimkewitsch  1891 
Phylum  Chaetognatha  Leuckart  1854 
Phylum  Entoprocta  Nitsche  1870 
Phylum  Bryozoa  Ehrenberg  1831 
Class  Gymnolaemata  Allman  1856 
Order  Cyclostomata  Busk  1852 
Suborder  Tubipora  Hagenow  1851 
Suborder  Cerioporina  Hagenow  1851 
Suborder  Ceramoporoidea  Bassler  1913 
Suborder  Hedrelloidea  Bassler  1934 
Order  Trepostomata  Ulrich  1883 
Order  Cryptostomata  Vine  1883 
Order  Ctenostoma  Busk  1852 

[9] 


Order  Cyclostomata  Busk  1852 
Suborder  Anasca  Levinsen  1909 
Suborder  Ascophora  Levinsen  1909 
Class  Phylactolaemata  Allman  1856 
Phylum  Brachiopoda  Cuvier  1802 
Class  Inarticulata  Huxley  1869  (Gastrocaulia  Thompson  1927) 
Order  Atremata  Beecher  1891 
Order  Neotremata  Beecher  1891 
Class  Articulata  Huxley  1869 

Order  *Paleotremata  Thompson  1927 
Order  Protremata  Beecher  1891 
Order  Telotremata  Beecher  1891 
Phylum  Phoronidea  E.  Shultz  1903  (Pygocaulia  Thompson  1927) 
Phylum  Pogonophora  Johansson  1937 
Phylum  Pterobranchia  Lankester  1877 
Phylum  Enteropneusta  Gegenbaur  1870 

Class  Balanoglossida  Delle  Chiaje  1820 
Phylum  Linguatula  Frolich  1789 
Phylum  Echinodermata  Laske  1778 
Subphylum  Pelmatozoa  Leuckart  1848 
Class  *Cystoidea  von  Buch  1844 
Order  *Amphoridea  Haeckel  1896 
Order  *Rhombifera  Zittel  1879 
Order  *Diploporita  Zittel  1879 
Order  *Aporita  Zittel  1879 
Order  *Edrioasteroida  Billings  1858 
Class  *Blastoidea  Say  1925 

Order  *Protoblastoida  Bather  1899 
Order  *Eublastoida  Bather  1899 
Class  Crinoidea  Miller  1921 

Order  *Camerata  Wachsmuth  &  Springer  1885 
Order  *Adunata  Bather  1899 
Order  Flexibilia  Zittel  1879 

Suborder  *Sagenocrinida  Springer  1926 
Suborder  *Taxocrinida  Springer  1926 
Order  *Inadunata  Wachsmuth  &  Springer  1885 

Suborder  *Larviformia  Wachsmuth  &  Springer  1885 
Suborder  *Fistulata  Wachsmuth  &  Springer  1885 
Order  Articulata  Aliller  1821 

Suborder  Bourgueticrinida  H.  L.  Clark  1939 
Suborder  Pentacrinida  Claus  1868 
Suborder  Holopida  Haeckel  1896 
Order  Comatulida  (Parks  1891)  A.  H.  Clark  1908 
Suborder  Oligophreata  A.  H.  Clark  1908 
Suborder  Macrophreata  A.  H.  Clark  1908 
Subphylum  Asterozoa  Leuckart  1848 
Class  Asteroidea  Burmeister  1837 
Order  Phanerozonea  Sladen  1889 
Order  Spinulosa  Perrier  1894 
Order  Forcipulata  Perrier  1894 
Suborder  Brisingina  Fisher  1928 
Suborder  Asteriadina  Fisher  1928 

[10] 


Class  Ophiuroidea  Norman  1865 
Subclass  *Aegophiuricia  Matsumoto  1915 
Order  *Lysoi)hiiirida  Matsumoto  1913 
Subclass  Myophiurida  Matsumoto  1915 
Order  *Opbiocystiida  H.  L.  Clark  1939 
Order  *Aganasterida  H.  L.  Clark  1939 
Order  Phrynophiurida  Matsumoto  1915 
Order  Laemophiurida  Matsumoto  1915 
Order  Gnathophiurida  Matsumoto  1915 
Order  Chilophiurida  Matsumoto  1915 
Subphylum  Echinozoa  Leuckart  1848 
Class  Echinoidea  Bronn  1860 
Order  *Bothriocidaroida  Duncan  1889 
Order  Cidaroida  Duncan  1889 
Order  Centrechinoida  Jackson  1912 
Suborder  Aulodonta  Jackson  1912 
Suborder  Stirodonta  Jackson  1912 
Suborder  Camerodonta  Jackson  1912 
Order  Exocycloida  Jackson  1912 
Suborder  Holectypina  Gregory  1900 
Suborder  Echinoneina  Hawkins  1925 
Suborder  Clypeastrina  Gregory  1900 
Suborder  Nucleolitina  Hawkins  1925 
Suborder  Cassidulina  Hawkins  1925 
Suborder  Urechinina  H.  L.  Clark  1925 
Suborder  Spatangina  Jackson  1912 
Order  Perischoechinoida  Jackson  1912 
Order  Echinocystoida  Jackson  1912 
Order  Perichoechinoida  Jackson  1912 
Class  Holothurioidea  Semper  1868 
Order  Dendrochirota  MacBride  1909 
Order  Elasipoda  Theele  1882 
Order  Aspidochirota  MacBride  1909 
Order  Molpadonia  Ekman  1925 
Order  Apoda  Lanipert  1885 
Phylum  Priapuloidea  Sedgwick  1898 
Phylum  Annelida  Lamarck  1809 
Class  Archiannelida  Hatschek  1893 
Class  Polychaeta  Grube  1850 
Order  Errantia  Audouin  &  Milne-Edwards  1832 
Order  Sedentaria  Lamarck  1818 
Order  Aplacophora  von  Ihreing  (see  page  17)  1878 
Order  *Miskoa  Walcott  1911 
Class  Myzostoma  Graff  1884 
Class  Oligochaeta  Grube  1850 
Order  Plesiopora  plesiotheca  Michaelsen  1930 
Order  Plesiopora  prosotheca  Michaelsen  1930 
Order  Prosopora  Michaelsen  1930 
Order  Opisthopora  Michaelsen  1930 
Class  Hirudinea  Lamarck  1818 

Order  Rhynchobdellida  Blanchard  1887 
Order  Gnathobdellida  Vaillant  1890 

[11] 


Order  Pharyngobdellida  Johansson  1913 
Phylum  Poeobioidea  Fisher  1946 
Phylum  Echiuroidea  Sedgwick  1898 
Class  Echiurida  Savigny  1817 
Order  Echiuroinea  Bock  1942 
Order  Xenopneusta  Fisher  1946 
Order  Heteromyota  Fisher  1946 
Class  Saccosomatida  Fisher  1946 
Phylum  Sipunculoidea  Sedgwick  1898 
Phylum  Onychophora  Grube  1851 
Phylum  Arthropoda  Siebold  &  Stannius  1845 
Subphylum  *Trilobita  Schlotheim  1820 
Class  *Hypoparia  Beecher  1897 
Class  *Opisthoparia  Beecher  1897 
Class  *Proparia  Beecher  1897 
Subphylum  Chelicerata  Dahl  1913 
Class  Merostomata  Woodward  1878 
Order  *Eurypterida  Woodward  1878 
Order  *Synxiphosura  Packard  1885 
Order  Xiphosura  Latreille  1806 
Class  Pycnogonida  Latreille  1804 
Order  Colossendeomorpha  Cole  1905 
Order  Nymphonomorpha  Pocock  1904 
Order  Ascorhynchomorpha  Pocock  1904 
Order  Pycnogonomorpha  Pocock  1904 
Class  Arachnida  Lamarck  1815 
Subclass  Latigastra  Petrunkevitch  1946 
Order  Scorpiones  Latreille  1810 
Order  Pseudoscorpiones  Latreille  1810 
Order  Opiliones  Sundevall  1833 
Order  *Architarbi  Petrunkevitch  1946 
Order  Acari  Leach  1817 

Suborder  Notostigmata  With  1903 
Suborder  Holothyroidea  Reuter  1909 
Suborder  Parasitiformes  Reuter  1909 
Suborder  Trombidiformes  Reuter  1909 
Suborder  Sarcoptiformes  Reuter  1909 
Suborder  Tetrapodili  Bremi  1872 
Subclass  Stethostomata  Petrunkevitch  1948 
Order  *Haptopoda  Pocock  1911 
Order  *Anthracomarti  Karsh  1882 
Subclass  Soluta  Petrunkevitch  1948 

Order  Trigonotarbi  Petrunkevitch  1948 
Subclass  Caulogastra  Pocock  1910 

Superorder  Latisterna  Petrunkevitch  1948 

Order  Palpigradi  Thorell  1888 

Superorder  Camarostomata  Petrunkevitch  1948 

Order  Schizomida  Ptrunkevitch  1913 

Order  Telyphonida 

Order  Kustarachnae  Petrunkevitch  1913 

Superorder  Labellata  Petrunkevitch  1948 

Order  Phrynichida  Petrunkevitch  1946 

[12] 


Order  Araneae  Lamarck  1818 

Superorder  CucuUifera  Pocock  1910 

Order  Ricinulei  Latreille  1802 

Superorder  Rostrata  Petrunkevitch  1948 

Order  Solifugae  Sundevall  1833 
Subphylum  Mandibulata  McLeay  1821 
Superclass  Crustacea  Pennant  1777 
Class  Eucrustacea  Kingsley  1894 
Subclass  Branchiopoda  Lamarck  1801 

Order  Anostraca  Sars  1867 

Order  Notostraca  Sars  1867 

Order  Conchostraca  Sars  1867 

Order  Cladocera  Latreille  1829 
Suborder  Calyptomera  Sars  1865 
Suborder  Gymnomera  Sars  1865 
Subclass  Ostracoda  Latreille  1802 

Order  Myodocopa  Sars  1866 

Order  Cladocopa  Sars  1866 

Order  Podocopa  Sars  1866 

Order  Platycopa  Sars  1866 
Subclass  Copepoda  Latreille  1831 

Order  Eucopepoda  Claus  1875 
Suborder  Calanoida  Dana  1852 
Suborder  Harpacticoida  Sars  1901 
Suborder  Cyclopoida  Burmeister  1834 
Suborder  Notodelphyoida  Sars  1903 
Suborder  Monstrilloida  Sars  1903 
Suborder  Caligoida  Claus  1885 
Suborder  Lernaeopodoida  Stebbing  1910 

Order  Branchiura  Burmeister  1834 
Suborder  Arguloida  Wilson  1932 
Subclass  Cirripedia  Burmeister  1834 

Order  Thoracica  Darwin  1854 

Suborder  Turrilepadomorpha  Clarke  1896 
Suborder  Lepadomorpha  Darwin  1852 
Suborder  Verrucomorpha  Darwin  1854 
Suborder  Balanomorpha  Darwin  1854 

Order  Ascothoracica  Lacaze-Duthiers  1883 

Order  Apoda  Darwin  1854 

Order  Rhizocephala  F.  Miiller  1862 
Subclass  Malacostraca  Latreille  1802 
Infraclass  Leptostraca  Claus  1880 

Division  Phyllocarida  Packard  1879 

Order  Nebaliacea  Clarke  1882 

Order  *Rhinocarina  Clarke  1882 

Order  *Ceratiocarina  Clarke  1882 

Order  *Hymenocarina  Clarke  1882 

Division  *Nahecarida  Broili  1928 
Infraclass  Eumalacostraca  Grobben  1892 

Division  Syncarida  Packard  1886 

Order  Anaspidacea  Thomson  1892 

Division  Peracarida  Caiman  1904 

[13] 


Order  Mysidacea  Boas  1883 
Order  Thermosbaenacea  Monod  1927 
Order  Cuniacea  Kr0yer  1846 
Order  Tanaidacea  Hansen  1895 
Order  Isopoda  Latreille  1817 
Suborder  Flabellifera  Sars  1882 
Suborder  Valvifera  Sars  1882 
Suborder  Asellota  Latreille  1806 
Suborder  Phreatoicidea  Stebbing  1903 
Suborder  Epicaridea  Latreille  1831 
Suborder  Oniscoidea  Sars  1882 
Order  Amphipoda  Latreille  1816 
Suborder  Gammaridea  Dana  1852 
Suborder  Hyperiidea  Dana  1852 
Suborder  Caprellidea  Dana  1852 
Suborder  Ingolfiellidea  Hansen  1903 
Division  Eucarida  Caiman  1904 
Order  Euphausiacea  Boas  1883 
Order  Decapoda  Latreille  1802 
Suborder  Natantia  Boas  1879 
Suborder  Reptantia  Boas  1879 
Division  Hoplocarida  Caiman  1904 
Order  Stomatopoda  Latreille  1817 
Superclass  Progoneata  Pocock  1893 
Class  Pauropoda  Lubbock  1866 
Order  Pauropoda  Lubbock  1866 
Suborder  Ectomorpha  Bagnall  1935 
Suborder  Endomorpha  Bagnall  1935 
Class  Symphyla  Ryder  1880 

Order  Symphyla  Ryder  1880 
Class  Diplopoda  Latreille  1802 
Subclass  Pselaphognatha  Latzel  1884 
Order  Ancyrotricha  Cook  1895 
Order  *Lophotricha  Cook  1895 
Subclass  Chilognatha  Leach  1814 

Superorder  Opisthandria  Verhoefif  1894 
Order  Limacomorpha  Pocock  1894 
Order  Oniscomorpha  Pocock  1887 
Suborder  Sphaerotheria  Brandt  1833 
Suborder  Glomeridia  Brandt  1833 
Superorder  Proterandria  Verhoeff  1894 
Order  Colobognatha  Brandt  1834 
Order  Nematophora  Verhoeff  1913 
Suborder  Chordeumoidea  Latzel  1884 
Suborder  Stemmiuloidea  Cook  1895 
Suborder  Striaroidea  Cook  1896 
Suborder  Lysiopetaloidea  Verhoeff  1900 
Order  Proterospermophora  Verhoeff  1913 
Suborder  Polydesmoidea  Attems  1914 
Suborder  Strongylosomoidea  Attems  1914 
Order  Opisthospermophora  Verhoeff  1913 
Suborder  Juloidea  Meinert  1868 

[14] 


Suborder  Spiroboloidea  Sylvestri  1896 
Suborder  Spirostreptoidea  Pocock  1894 
Suborder  Cambaloidea  BoUnian  1893 
Superclass  Opisthogoneata  Pocock  1893 
Class  Chilopoda  Latreille  1802 
Subclass  Plcurostigma  Pocock  1902 
Superorder  Epimorplia  Haase  1880 
Order  Geophilomorpha  Leach  1814 
Order  Scolopendroniorpha  Newport  1844 
Superorder  Anamorpha  Haase  1880 
Order  Lithobiomorpha  Verhoeff  1907 
Order  Craterostigma  Pocock  1902 
Subclass  Notostigma  Pocock  1902 

Order  Scutigcromorpha  Pocock  1895 
Class  Insecta  Linnaeus  1758 
Subclass  Synaptera  Packard  1883 
Order  Collembola  Lubbock  1871 
Order  Protura  Silvestri  1907 
Order  Entotrophi  Grassi  1888 
Order  Thysanura  Latreille  1796 
Subclass  Pterygota  (Brauer  1885)  Lang  1889 
Order  Odonata  Fabricius  1793 
Order  Plectoptera  Packard  1886 
Order  Orthoptera  Latreille  1796 
Order  Isoptera  Brulle  1832 
Order  Dermaptera  Leach  1815 
Order  Plecoptera  Burmeister  1835 
Order  Embioptera  Hagen  1861  (Shipley  1904) 
Order  Corrodentia  Burmeister  1839 
Order  Zoraptera  Silvestri  1913 
Order  Mallophaga  Nitsch  1818 
Order  Anoplura  Leach  1815 
Order  Hemiptera  Linnaeus  1758 

Suborder  Heteroptera  Latreille  1825 
Suborder  Homoptera  Latreille  1817 
Order  Thysanoptera  Haliday  1836 
Order  Neuroptera  Linnaeus  1758 
Order  Mecoptera  Packard  1886 
Order  Diptera  Linnaeus  1758 

Suborder  Orthorrhapha  Brauer  1863 
Suborder  Cyclorrhapha  Brauer  1863 
Order  Siphonaptera  Latreille  1825 
Order  Trichoptera  Kirby  1813 
Order  Lepidoptera  Linnaeus  1758 
Suborder  Jugatae  Comstock  1892 
Suborder  Frenatae  Comstock  1892 
Suborder  Rhopalocera  Dumeril  1806 
Order  Megaloptera  Latreille  1802 
Order  Raphidiodea  Burmeister  1835 
Order  Coleoptera  Linnaeus  1758 

Suborder  Adephaga  Leconte  &  Horn  1883 
Suborder  Polyphaga  Emery  1885 

[15] 


Order  Strepsiptera  Kirby  1813 
Order  Hymenoptera  Linnaeus  1758 

Suborder  Chalastogastra  Konow  1897 

Suborder  CHstogastra  Konow  1905 
Order  *Palaeodictyoptera  Goldenberg  1877 
Order  *Megasecoptera  Brongniart  1885 
Order  *Protephemerida  Handlirsch  1906 
Order  *Protodonata  Brongniart  1885 
Order  *Protoperlaria  Tillyard  1928 
Order  *Protelytroptera  Tillyard  1931 
Order  *Caloneurodea  Martynov  1938 
Order  *Glosselytrodea  Martynov  1938 
Order  *Protorthoptera  Handlirsch  1906 
Order  *Protohemiptera  Handlirsch  1906 

The  following  extinct  orders  of  insects  have  also  been  described,  but,  according  to 
F.  M.  Carpenter,  probably  do  not  "deserve  recognition" : 
Order  *Protoblattoidea  Handlirsch  1906 
Order  *Hadentomoidea  Handlirsch  1906 
Order  *Sypharopteroidea  Handlirsch  1906 
Order  *Mixotermitoidea  Handlirsch  1906 
Order  *Reculoidea  Handlirsch  1906 
Order  *Hapalopteroidea  Handlirsch  1906 
Order  *Protomecoptera  Tillyard  1917 
Order  *Paratrichoptera  Tillyard  1919 
Order  *Paramecoptera  Tillyard  1919 
Order  *Synarmogoidea  Handlirsch  1919 
Order  *Diaphanopteroidea  Handlirsch  1919 
Order  *Aeroplanoptera  Tillyard  1923 
Order  *Protohymenoptera  Tillyard  1923 
Order  *Protocoleoptera  Tillyard  1923 
Order  *Miomoptera  Martynov  1927 
Order  *Pruvostitoptera  Zalessky  1929 
Order  *Meganisoptera  Martynov  1931 
Order  *Permodonata  Zalessky  1931 
Order  *Archodonata  Martynov  1932 
Order  *Protodiptera  Tillyard  1935 
Order  *Hemipsocoptera  Zalessky  1937 
Order  *Cnemidolestoidea  Handlirsch  1937 
Order  *Paraplecoptera  Martynov  1938 
Order  *Archaehymenoptera  Haupt  1941 
Order  *Palaeohymenoptera  Haupt  1941 
Order  *Protocicadida  Haupt  1941 
Order  *Protofulgorida  Haupt  1941 
Order  *Hemiodonata  Zalessky  1943 
Order  *Anisaxia  Forbes  1943 
Order  *Perielytrodea  Zalessky  1943 
Order  *Permodictyoptera  Zalessky  1944 
Order  *Aphelophlebia  Pierce  1945 
Phylum  MoUusca  Linnaeus  1758 

Class  Crepipoda  Goldfuss  1820  ( Polyplacophora  Gray  1821) 
Order  Eoplacophora  Pilsbry  1892 

[16] 


Order  Aplacophora  von  Ihering  1877 
Suborder  Chaetodermatina  Simroth  1893 
Suborder  Neomeniina  Simroth  1893 
Order  Mesoplacophora  Pilsbry  1892 
Order  Isoplacophora  Cotton  &  Weeding  1939 
Order  Teleoplacophora  Pilsbry  1892 
Class  Gastropoda  Cuvier  1798 
Subclass  Prosobranchia  Milne-Edwards  1848 

Order  Archaeogastropoda  Thiele  1925  (Aspidobranchia  Pelseneer  1890) 
Order  Mesogastropoda  Thiele  1925 

Order  Neogastropoda  Wenz  1938  (Stenoglossa  Bouvier  1887) 
Subclass  Opisthobranchia  Milne-Edwards  1846 
Order  Tectibranchia  Cuvier  1812  (Pleurocoela  Thiele  1931) 
Order  Pteropoda  Cuvier  1804 

Suborder  Thecosomata  Blainville  1824 
Suborder  Gymnosomata  Blainville  1824 
Order  Sacoglossa  von  Ihering  1876 
Order  Acoela  Thiele  1931 

Suborder  Notaspidea  Fischer  1883 
Suborder  Nudibranchia  Cuvier  1817 
Subclass  Pulmonata  Cuvier  1817 
Order  Basommatophora  A.  Schmidt  1895 
Order  Stylommatophora  A.  Schmidt  1895 
Class  Scaphopoda  Bronn  1862 
Class  Pelecypoda  Goldfuss  1820 
Subclass  Prionodesmacea  Dall  1889 
Order  Palaeoconcha  Neumayr  1883 
Order  Taxodonta  Neumayr  1883 
Order  Schizodonta  Steinmann  1890 
Order  Dysodonta  Neumayr  1883 
Subclass  Anomalodesmacea  Dall  1889 
Order  Anomalobranchia  Iredale  1939 
Order  Septibranchia  Perrier  1897 
Subclass  Teleodesmacea  Dall  1889 
Order  *Pantodonta  Dall  1895 
Order  Diogenodonta  Dall  1895 
Order  Cyclodonta  Dall  1895 
Order  Teleodonta  Dall  1895 
Order  Asthenodonta  Dall  1895 
Class  Cephalopoda  Cuvier  1798 
Order  Tetrabranchia  Owen  1832 
Suborder  Nautiloidea  Lamarck  1809 
Suborder  *Ammonitoidea  Haan  1825 
Order  Dibranchia  Owen  1832 

Suborder  Vampyromorpha  Robson  1929 
Suborder  Decapoda  Leach  1817 
Suborder  Octopoda  Hasselquist  1850 
Phylum  Chordata  Haeckel  1874 
Subphylum  Tunicata  Lamarck  1816  (Urochordata) 
Class  Ascidiacea  Blainville  1825 
Order  Stolidobranchiata  Lahille  1887 
Order  Aspiraculata  Seeliger  1909 

[17] 


Order  Phlebobranchiata  Lahille  1887 
Order  Aplousobranchiata  Lahille  1887 
Class  Larvacea  Herdman  1882 
Class  Thaliacea  Van  der  Hoeven  1850 
Subclass  Myosomata  Garstang  1895 
Order  Salpida  Jones  1848 
Order  DoHolida  Keferstein  1862 
Subclass  Pyrosomata  Garstang  1895 
Order  Pyrosomatida  Jones  1848 
Division  Acrania  Hackel  1866  (Cephalochorda) 
Subphylum  Leptocardii  J.  Miiller  1845 
Class  Amphioxi  J.  Miiller  1844 
Order  Amphioxi  J.  Miiller  1844 
Division  Craniata  Lankester  1877  (Vertebrata) 
Subdivision  Anamnia 

Superclass  Pisces  Linnaeus  1758 
Class  Agnatha 

Subclass  Cephalaspidomorphi 
Order  *Osteostraci  Lankester  1868 
Order  *Anaspida  Traquair  1899 
Order  Cyclostomata 

Suborder  Myxinoidea  (Hyperotriti  Miiller  1844) 
Suborder  Petromyzontia  (Hyperoustii  Mulier  1844) 
Subclass  *Pteraspidomorphi 
Order  *Heterostraci  Lankaster  1868 
Order  *Coelolepida 
Class  *Placodermi  McCoy  1848 
Order  *Acanthodii  Owen  1860 
Order  *Arthrodira  Cope  1891 
Suborder  *Euarthrodira 
Infraorder  *Arctolepida 
Infraorder  *Brachythoraci 
Suborder  *Ptyctodontida  Woodward  1891 
Suborder  *Phyllolepida 
Order  *Macropetalichthyida 
Order  *Antiarchi  Cope  1885 
Order  *Stegoselachii 
Suborder  *Stensioellida 
Suborder  *Rhenanida 
Order  *Palaeospondyloidea  Sollas  1903 
Class  Chondrichthyes  J.  Mulier  1846 
Subclass  Elasmobranchii  Bonaparte  1832 
Order  *Cladoselachii  Dean  1894 
Order  Selachii  Cuvier  1829 
Suborder  *Hybodontoidea 
Suborder  Heterodontoidea 
Suborder  Notidanoidea 
Suborder  Galeoidea 
Suborder  Squaloidea 
Order  Batoidea  Gunther  1870 
Order  *Pleuracanthodii  Cope  1889 
Subclass  Holocephali  Bonaparte  1832 

[18] 


Order  *Bradyodonti  Woodward  1892 
Order  Chimaerae 
Class  Osteichthyes 
Subclass  Actinopterygii  Cuvier  1828 

Superorder  Chondrostei  G.  Miiller  1846  (Ganoids) 
Order  *Palaeoniscoidea  Bridge  1879 
Order  Polypterini  Bonaparte  1837 
Order  Acipenseroidei  Agassiz  1843 
Order  *Subholostei 
Superorder  Holostei  J.  Miiller  1846  (Ganoids) 
Order  *Semionotoidea  Woodward  1890 
Order  *Pycnodontoidea 
Order  *Aspidorhynchoidea 
Order  Amioidea  Huxley  1861 
Order  *Pholidophoroidea  Woodward  1890 
Superorder  Teleostei  Owen  1847 
Order  Isospondyli  Cope  1872 

Suborder  Clupeoidea 

Suborder  Salmonoidea 

Suborder  Opisthoproctoidea 

Suborder  Osteoglossoidea 

Suborder  Stomiatoidea 

Suborder  Gonorhynchoidea 
Order  Ostariophysi  Bonaparte  1809 

Suborder  Cyprinoidea 

Suborder  Siluroidea 
Order  Apodes  Rafinesque  1815 
Order  Heteromi  Gill 
Order  Mesichthyes 

Suborder  Haplomi  Cope  1872 

Suborder  Iniomi  Gill  1894 

Suborder  Cyprinodontes  Agassiz 

Suborder  Synentognathi  Cope  1872 

Suborder  Thoracostei 

Suborder  Salmopercae 
Order  Acanthopterygii  Cuvier  1828 

Suborder  Berycoidea 

Suborder  Zeoidea 

Suborder  Percoidea 

Suborder  Carangoidea 

Suborder  Scombroidea 

Suborder  Trachinoidea 

Suborder  Blennioidea 

Suborder  Anacanthini  Miiller  1846 

Suborder  Chaetodontoidea 

Suborder  Plectognathi  Cuvier  1817 

Suborder  Hetrosomata  Bonaparte  1846 

Suborder  Scorpaenoidea 

Suborder  Batrachoidea 

Suborder  Pediculati  Gill  1865 

Suborder  Gobioidea 

Suborder  Anabantoidea 

[19] 


Suborder  Mugiloidea 

Suborder  Polynemoidea 

Suborder  Ammodytoidea 

Suborder  Echeneoidea 

Suborder  Xenopterygii  Gill  1894 

Suborder  Allotriognathi  Regan  1907 

Suborder  Opisthomi  Cope  1870 

Suborder  Synbranchii 
Subclass  Amphibioidei  Hubbs  1911 
Order  Crossopterygii  Huxley  1861  (Ganoids) 

Suborder  *Rhipidistia  Cope  1887 

Suborder  Coelacanthini  Agassiz  1843 
Order  Dipnoi  J.  Miiller  1846 
Superclass  Tetrapoda  Williston 
Class  Amphibia  Linnaeus  1858 

Superorder  *Labyrinthodontia  Huxley  1871 
Order  *Ichthyostegalia  Save-Soderberg  1832 
Order  *Rhachitomia  Cope  1880 

Suborder  *Lexommoidea 

Suborder  *Edopsoidea 

Suborder  *Eryopsoidea 

Suborder  *Neorhachitomi 
Order  *Stereospondyli  Fraas  1889 
Order  ^Embolomeri  Cope  1880 
Order  *Seymouriamorpha 
Superorder  Salientia  Laurenti  1768 
Order  *Eoanura 
Order  *Proanura 
Order  Anura 

Suborder  Amphicoela  Nobel  1931 

Suborder  Opisthocoela  Nicholls  1916 

Suborder  Anomocoela  Nicholls  1916 

Suborder  Procela  Nicholls  1916 

Suborder  Diplasiocoela  Nicholls  1916 
Subclass  *Lepospondyli  Zittel  1887 
Order  *Aistopoda  Miall  1890 
Order  *Nectridia  Miall  1890 
Order  *Microsauria 
Order  Urodela  (Caudata  Oppel  1811) 

Suborder  Cryptobranchoidea  Dana  1922 

Suborder  Amblystomoidea  Noble  1931 

Suborder  Salamandriodea  Cope  1890 

Suborder  Proteidea  Cope  1866 
Order  Apoda  Oppel  1911 
Class  Reptilia  Laurenti  1768 
Subclass  Anapsida  Osborn  1918 
Order  *Cotylosauria  Cope  1880 

Suborder  *Captorhinomorpha  Watson  1917 

Suborder  *Diadectomorpha  Case  1911 
Order  Chelonia  Owen  1842  (Testudinata  Oppel  1811) 

Suborder  *Eunotosauria  Watson  1914 

Suborder  *Amphichelydia  Lydekker  1889 

Suborder  Plcurodira  Cope  1869 

[20] 


Suborder  Cryptodira  Cope  1869 
Subclass  *Ichthyopterygia  Owen  1860 
Subclass  *Synaptosauria  Williston  1912 

Order  *Protorosauria  Seeley  1871 

Order  *Sauropterygia  Owen  1859 
Suborder  *Nothsauria  Deecke  1866 
Suborder  *Plesiosauria  Andrews  1895 
Suborder  *Placodontia  Meyer  1863 
Subclass  Lepidosauria  Zittel  1890 

Order  *Eosuchia  Broom  1814 

Suborder  *Younginifonnes  Romer  1945 
Suborder  *Choristodera  Cope  1876 
Suborder  *Thalattosauria  Merriam  1904 
Suborder  *Acrosauria  Kuhn  1937 

Order  Rliynchocephalia  Gunther  1886 

Order  Squamata  Oppel  1811 

Suborder  Lacertilia  Cuvier  1834 
Suborder  Serpentes  Linnaeus  1758 
Subclass  *Archosauria  Cope  1891 

Order  *Thecodontia  Owen  1860 
Suborder  *Pseudosuchia  Cope  1891 
Suborder  *Pliytosauria  Lydekker  1888 

Order  Crocodilia  Wagler  1830 

Suborder  *Protosuchia  Mook  1934 
Suborder  *Sebecosuchia  Simpson  1937 
Suborder  *Mesosuchia  Mook  1934 
Suborder  Eusuchia  Mook  1934 

Order  *Pterosauri  Kaup  1834 

Suborder  *Rhamphorhynchoidea  Abel  1919 
Suborder  *Pterodactyloidea  Seeley  1866 

Order  *Saurischia  Seeley  1888 
Suborder  *Theropoda  Marsb  1881 
Suborder  *Sauropoda  Marsh  1878 

Order  *Ornithischia  Seeley  1888 

Suborder  *Ornithopoda  Cope  1869 
Suborder  *Ankylosauria  Brown  1908 
Suborder  *Ceratopsia  Zittel  1890 
Subclass  *Synapsida  Osborn  1903 

Order  *Pelycosauria  Cope  1878 

Suborder  *Ophiacodontia  Romer  &  Price  1940 
Suborder  *Sphenacodontia  Romer  &  Price  1940 
Suborder  *Edaphosauria  Romer  &  Price  1940 

Order  *Therapsida  Broom  1905 

Suborder  *Dinocephalia  Broom  1914 
Suborder  *Dicynodontia  Owen  1859 
Suborder  *Theriodontia  Owen  1859 

Order  *Ictidosauria  Broom  1912 
Class  Aves  Linnaeus  1758 
Subclass  *Archaeornithes  Cope  1898 

Order  *Archaeopterygiformes  Fiirbringer 
Subclass  Neornithes  Reynolds  1897 

Superorder  *Odontognathae  Marsh  1873 

Order  *Hesperornithiformes  Marsh  1872 

[21] 


Order  *Ichthyornithiformes  Marsh  1873 
Superorder  Palaeognathae  Pycraft  1910 
Order  *Caenagnathiformes 
Order  Struthioniformes  Haeckel  1866 
Order  Rheiformes  Newton  1884 
Order  Casuariiformes  Sclater  1880 
Order  *Dinornithifornies  Stejneger  1884 
Order  *Aepyornithiformes  Newton  1884 
Order  Apterygiformes  Wetmore  &  Miller  1936 
Order  Tinamiformes  Gray  1840 
Superorder  Impennes  Illiger  1811 
Order  Sphenisciformes  Viellot  1818 
Superorder  Neognathae  Pycraft  1896 
Order  Gaviiformes  Brandt  1839 
Order  Colymbifornies  Bonaparte  1852 
Order  Procellariiformes  Wagner  1843 
Order  Pelecaniformes  Stejneger  1885 
Suborder  Phaethontes  Gray  1840 
Suborder  Pelecani  Owen  1856 
Suborder  Fregatae  Vigors  1830 
Suborder  Odontopteryges  Lydekker  1892 
Order  Ciconiiformes  Owen  1832 
Suborder  Ardeinae  Wagler  1830 
Suborder  Balaenicipites  Gould  1851 
Suborder  Ciconiae  Bonaparte  1834 
Suborder  Phoenicopteri  Bonaparte  1831 
Order  Anseriformes  Swainson  1837 
Suborder  Anhimae  Brisson  1760 
Suborder  Anseres  Linnaeus  1735 
Order  Falconiformes  Naumann  1822 
Suborder  Cathartae  Lafresnaye  1829 
Suborder  Falcones  Naumann  1822 
Order  Galliformes  Latreille  1825 
Suborder  Galli  Dumeril  1806 
Suborder  Opisthocomi  Gray  1840 
Order  Gruiformes  Stejneger  1885 
Suborder  Mesoenatides  Lowe  1924 
Suborder  Turnices  Gray  1840 
Suborder  Grues  Owen  1856 
Suborder  Heliornithes  Gray  1840 
Suborder  Rhynocheti  Gould  1837 
Suborder  Eurypygae  Illiger  1811 
Suborder  *Phororhaci  Lydekker  1893 
Suborder  Cariamae  Brisson  1760 
Suborder  Otides  Wagler  1830 
Order  *Diatrymiformes  Cope  1877 
Order  Charadriiformes  Fiirbinger  1888 
Suborder  Charadrii  Huxley  1867 
Suborder  Lari  Bonaparte  1857 
Suborder  Alcae  Stejneger  1885 
Order  Columbiformes  Bonaparte  1853 
Suborder  Pterocletes  Knowlton  1909 
Suborder  Columbae  Bonaparte  1857 

[221 


Order  Psittaciformes  Illiger  1811 
Order  Cuculiformes  Stejneger  1884 
Suborder  Musophagi  Sharpe  1891 
Suborder  Cuculi  A.  O.  U.  1886 
Order  Strigiformes  Wagler  1830 
Order  Caprimulgiformes  Stejneger  1885 
Suborder  Steatornithes  Bonaparte  1850 
Suborder  Caprimulgi  Bonaparte  1840 
Order  Apodiformes  Lucas  1889 
Suborder  Apodi  Peters  1940 
Suborder  Trochili  Lesson  1931 
Order  Coliiformes  Swainson  1837 
Order  Trogoniformes  Swainson  1831 
Order  Coraciiformes  Seebohm  1880 
Suborder  Alcedines  Bonaparte  1831 
Suborder  Meropes  Bonaparte  1831 
Suborder  Coracii  Gray  1845 
Suborder  Bucerotes 
Order  Piciformes  Sharpe  1891 

Suborder  Galbulae  Fiirbinger  1888 
Suborder  Pici  Wagler  1830 
Order  Passeriformes  Nitzsch  1829 
Suborder  Eurylaimi  Gadow  1893 
Suborder  Tyranni  Swainson  1837 
Suborder  Menurae  Stejneger  1885 
Suborder  Passeres  Linnaeus  1758 
Class  Mammalia  Linnaeus  1758 
Subclass  Prototheria  Gill  1872 

Order  Monotremata  Bonaparte  1838 
Subclass  *Allotheria  Marsh  1880 
Order  *Multituberculata  Cope  1884 
Order  *Triconodonta  Osborn  1888 
Subclass  Theria  Parker  &  Haswell  1897 
Infraclass  *Pantotheria  Simpson  1929 
Order  *Pantotheria  Marsh  1880 
Order  *Symmetrodonta  Simpson  1925 
Infraclass  Metatheria  Huxley  1880 

Order  Marsupialia  Illiger  1811 
Infraclass  Eutheria  Gill  1872 
Cohort  Unguiculata  Linnaeus  1766 
Order  Insectivora  Bowdich  1821 
Order  Dermoptera  Illiger  1811 
Order  Chiroptera  Blumenbach  1779 
Suborder  Megachiroptera  Dobson  1875 
Suborder  Microchiroptera  Dobson  1875 
Order  Primates  Linnaeus  1758 
Suborder  Prosimii  Illiger  1811 
Suborder  Anthropoidea  Mivart  1864 
Order  *Tillodontia  Marsh  1875 
Order  *Taeniodontia  Cope  1876 
Order  Edentata  Cuvier  1798 

Suborder  *Palaeanodonta  Matthew  1918 
Suborder  Xenarthra  Cope  1889 

[23] 


Order  Pholidota  Weber  1904 
Cohort  Glires  Linnaeus  1758 

Order  Lagomorpha  Brandt  1855 

Order  Rodentia  Bowdich  1821 

Suborder  Sciuromorpha  Brandt  1855 
Suborder  Myomorpha  Brandt  1855 
Suborder  Hystricomorpha  Brandt  1855 
Cohort  Mutica  Linnaeus  1766 

Order  Cetacea  Brisson  1762 

Suborder  *Archaeoceti  Fowler  1883 
Suborder  Odontoceti  Fowler  1867 
Suborder  Mysticeti  Fowler  1864 
Cohort  Ferungulata  Simpson  1945 

Superorder  Ferae  Linnaeus  1758 

Order  Carnivora  Bowdich  1821 
Suborder  *Creodonta  Cope  1875 
Suborder  Fissipeda  Blumenbach  1791 
Suborder  Pinnipedia  Illiger  1811 

Superorder  *Protungulata  Weber  1904 

Order  *Condylarthra  Cope  1881 

Order  *Litopterna  Ameghino  1889 

Order  *Notoungulata  Roth  1903 

Suborder  *Notioprogonia  Simpson  1934 
Suborder  *Toxodonta  Scott  1904 
Suborder  *Typotheria  Zittel  1892 
Suborder  *Hegetotheria  Simpson  1945 

Order  *Astrapotheria  Lydekker  1894 

Suborder  *Trigonostylopoidea  Simpson  1934 
Suborder  *Astrapotherioidea  Ameghino  1894 

Order  Tubulidentata  Huxley  1872 

Superorder  Paenungulata  Simpson  1945 

Order  *Pantodonta  Cope  1873 

Order  *Dinocerata  Marsh  1873 

Order  *Pyrotheria  Ameghino  1895 

Order  Proboscidea  Illiger  1811 

Suborder  *Moeritherioidea  Osborn  1921 
Suborder  Elephantoidea  Osborn  1921 
Suborder  *D€inotherioidea  Osborn  1921 
Suborder  *Barytherioidea  Simpson  1945 

Order  *Embrithopoda  Andrews  1906 

Order  Hyracoidea  Huxley  1869 

Order  Sirenia  Illiger  1811 

Suborder  Trichechiformes  Hay  1923 
Suborder  *Desniostyliformes  Hay  1923 

Superorder  Mesaxonia  Marsh  1884 

Order  Perissodactyla  Owen  1848 
Suborder  Hippomorpha  Wood  1937 
Suborder  Ceratomorpha  Wood  1937 

Superorder  Paraxonia  Marsh  1884 

Order  Artiodactyla  Owen  1848 
Suborder  Suiformes  Jaeckel  1911 
Suborder  Tylopoda  Illiger  1811 
Suborder  Ruminantia  Scopoli  1777 

[24] 


Price  25c 

Zoology  Department 

Duke  University 

Durham,  N.  C,  U.  S.  A.